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Equation 1 · Part 3 · One Gene, Forty Eyes: What Evo-Devo Did to the Convergence Story

Symbol V

h2×i×V=0.50×0.01×0.01=0.00005h^2 \times i \times V = 0.50 \times 0.01 \times 0.01 = 0.00005
VV

What this part means

V is part of the quantity the equation computes from the expression on the right.

Its job in the formula

V is part of the quantity the equation computes from the expression on the right.

The passage around this formula

To convert steps into generations, Nilsson and Pelger applied the standard quantitative-genetics equation for the response to selection on a continuous trait, deliberately choosing pessimistic values for every free parameter: a heritability of 0.50, a selection intensity of 0.01, and a coefficient of phenotypic variation of 0.01, values the authors describe as common or deliberately conservative rather than favourable to a fast result [ 4 ] . Those choices give a fractional improvement of only 0.005 percent of the trait mean per generation. Compounding that per-generation rate to reach the full 1,829 steps of one-percent change — equivalent to a single structure lengthening by a factor of…

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A variable is a named place for a value. Its letter is a local label: x can mean position in one formula and a data point in another.

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