Equation 33 · Part 1 · Evolutionary Biology and Ecology in Practice: An Advanced Technical Guide
Symbol N_e
What this part means
the downward in inferred.
Its job in the formula
is a part of this expression. Its role is fixed by the surrounding article and by the operations shown in the formula.
Full expression→Symbol N_e→Article meaning
Where the article explains it
Physically linked loci must be excluded or flagged , because the method’s entire logic depends on treating background disequilibrium as drift-generated between unlinked loci; loci close together on the same chromosome carry disequilibrium from physical linkage that has nothing to do with and will bias the estimate upward in apparent LD, downward in inferred .
The passage around this formula
What this means for study design , working backward from the estimator: . The genetic estimate and the demographic estimate from workflow one are not redundant. A mark-recapture study can show a stable or growing census count while an LD-based estimate reveals a much smaller effective size — a mismatch that is itself diagnostic, typically pointing to high variance in reproductive success (a few individuals contributing disproportionately to the next generation) or a historical bottleneck whose genetic signature outlasts the demographic recovery.
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Sources cited in the article section
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