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Equation 7 · Part 2 · Einstein's Random Walk and the Mathematics of Genetic Drift

Symbol R

D=RTN⋅16πkP,D = \frac{RT}{N}\cdot\frac{1}{6\pi k P},
RR

What this part means

the gas constant.

Its job in the formula

R occurs above the fraction bar. The numerator is divided by the entire denominator below it.

Where the article explains it

where R is the gas constant, T the absolute temperature, and N Avogadro’s number [ 1 ] .

The passage around this formula

with the diffusion constant D fixed by measurable quantities. Einstein derived it explicitly, from the balance between the drag on a sphere of radius P in a fluid of viscosity k and the osmotic pressure driving diffusion, as D=RTN⋅16πkPD = \frac{RT}{N}\cdot\frac{1}{6\pi k P}. where R is the gas constant, T the absolute temperature, and N Avogadro’s number [ 1 ] . That last dependency was the trap door: if D could be read off a microscope by timing how far a particle wandered, and every other term in the equation was already known independently, then watching one particle jiggle became a way to count atoms.

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Learn the underlying idea

A variable is a named place for a value. Its letter is a local label: x can mean position in one formula and a data point in another.

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Sources cited in the surrounding passage

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