Symbol w_open
pen is part of the quantity the equation computes from the expression on the right.
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Why this formula appears here
which approaches a fixed value of 1 + a/b as t grows without bound, and . for an exponent between 0 and 1, which keeps rising — more and more slowly, but without any asymptote — for every value of t . Wiser and colleagues derived the second form theoretically from a model combining clonal interference with diminishing-returns epistasis among beneficial mutations, and reported that across 50,000 generations of real data, “mean fitness appears to increase without bound, consistent with a power law” [ 2 ] . The decelerating rate that Lenski’s 1991 paper had already observed over the first 2,000 generations turns out to be compatible with no ceiling at all: a…
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Symbol t
t is an argument of the function-like quantity on the left; its role is set by that function’s stated inputs.
Read this term in its guide →Symbol a
a is one of the signed contributions combined to compute the quantity on the left.
Read this term in its guide →Symbol t^β
t^β is one of the signed contributions combined to compute the quantity on the left.
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Read it with the definitions, units, and assumptions supplied by the article.
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Published contexts (1)
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Equation 4 · Evolutionary Biology
Seventy-Five Thousand Generations of E. coli, and Counting
This equation states an equality: the expressions on both sides have the same value under the article’s assumptions.
which approaches a fixed value of 1 + a/b as t grows without bound, and . for an exponent between 0 and 1, which keeps rising — more and more slowly, but without any asymptote — for every value of t . Wiser and colleagues derived the second form theoretically from a model combining clonal interference with diminishing-returns epistasis among beneficial mutations, and reported that across 50,000 generations of real data, “mean fitness appears to increase without bound, consistent with a power law” [ 2 ] . The decelerating rate that Lenski’s 1991 paper had already observed over the first 2,000 generations turns out to be compatible with no ceiling at all: a…
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