Equation 4 · Part 1 · Seventy-Five Thousand Generations of E. coli, and Counting
Symbol w_open
What this part means
pen is part of the quantity the equation computes from the expression on the right.
Its job in the formula
pen is part of the quantity the equation computes from the expression on the right.
Full expression→Symbol w_open→Article meaning
The passage around this formula
which approaches a fixed value of 1 + a/b as t grows without bound, and . for an exponent between 0 and 1, which keeps rising — more and more slowly, but without any asymptote — for every value of t . Wiser and colleagues derived the second form theoretically from a model combining clonal interference with diminishing-returns epistasis among beneficial mutations, and reported that across 50,000 generations of real data, “mean fitness appears to increase without bound, consistent with a power law” [ 2 ] . The decelerating rate that Lenski’s 1991 paper had already observed over the first 2,000 generations turns out to be compatible with no ceiling at all: a…
Learn the underlying idea
A subscript is a label attached below a symbol. It often selects a time step, component, category, or member of a sequence.
Open the illustrated subscripts: which member of a family? guide →
See this notation across published equations →
Sources cited in the surrounding passage
These citations provide research context; check each source for the exact claim it supports.