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Published equation contexts

∂ϕ(p,t)∂t=12 ∂2∂p2[V(p) ϕ(p,t)],V(p)=p(1−p)2N(Kimura, 1955/1962)\frac{\partial \phi(p,t)}{\partial t} = \frac{1}{2}\,\frac{\partial^{2}}{\partial p^{2}}\Big[V(p)\,\phi(p,t)\Big], \quad V(p) = \frac{p(1-p)}{2N} \qquad \text{(Kimura, 1955/1962)}

Why this formula appears here

Motoo Kimura, at Japan’s National Institute of Genetics, made the borrowing exact rather than approximate. His 1955 paper in the Proceedings of the National Academy of Sciences solved the full time-dependent diffusion equation derived from the Wright-Fisher model, recovering not just the eventual resting distribution of an allele’s frequency but its whole transient shape as it evolves generation by generation [ 6 ] . His 1962 paper in Genetics then posed the general fixation problem as a diffusion boundary-value equation and named the lineage precisely: the question of a mutant gene’s ultimate fate was “first treated quantitatively by Fisher (1922),” with “equivalent results” reached…

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NN

Symbol N

N occurs below the fraction bar. The quantity above the bar is divided by this expression; zero is excluded as a denominator.

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∂p2\partial p^{2}

Denominator: partial p^2

The complete quantity below the fraction bar; it must be nonzero for this division.

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How to interpret it

With a fixed numerator, increasing a nonzero denominator reduces the fraction. Read it with the definitions, units, and assumptions supplied by the article.

Research cited beside this formula

Published contexts (1)

A symbol can carry a different meaning in another article. Each occurrence keeps its own guide and term definitions.

∂ϕ(p,t)∂t=12 ∂2∂p2[V(p) ϕ(p,t)],V(p)=p(1−p)2N(Kimura, 1955/1962)\frac{\partial \phi(p,t)}{\partial t} = \frac{1}{2}\,\frac{\partial^{2}}{\partial p^{2}}\Big[V(p)\,\phi(p,t)\Big], \quad V(p) = \frac{p(1-p)}{2N} \qquad \text{(Kimura, 1955/1962)}

Equation 21 · Einstein & Evolution

Einstein's Random Walk and the Mathematics of Genetic Drift

This equation states an equality: the expressions on both sides have the same value under the article’s assumptions.

Motoo Kimura, at Japan’s National Institute of Genetics, made the borrowing exact rather than approximate. His 1955 paper in the Proceedings of the National Academy of Sciences solved the full time-dependent diffusion equation derived from the Wright-Fisher model, recovering not just the eventual resting distribution of an allele’s frequency but its whole transient shape as it evolves generation by generation [ 6 ] . His 1962 paper in Genetics then posed the general fixation problem as a diffusion boundary-value equation and named the lineage precisely: the question of a mutant gene’s ultimate fate was “first treated quantitatively by Fisher (1922),” with “equivalent results” reached…

Meanings in this article

  • pp: the starting from frequency.
  • p2p^{2}: the square of p; the starting from frequency.
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