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Equation 31 · Part 1 · Evolutionary Biology and Ecology in Practice: An Advanced Technical Guide

Symbol N_e

NeN_e
NeN_e

What this part means

the downward in inferred.

Its job in the formula

NeN_e is a part of this expression. Its role is fixed by the surrounding article and by the operations shown in the formula.

Where the article explains it

Physically linked loci must be excluded or flagged , because the method’s entire logic depends on treating background disequilibrium as drift-generated between unlinked loci; loci close together on the same chromosome carry disequilibrium from physical linkage that has nothing to do with NeN_e and will bias the estimate upward in apparent LD, downward in inferred NeN_e .

The passage around this formula

What this means for study design , working backward from the estimator: NeN_e. The genetic estimate and the demographic estimate from workflow one are not redundant. A mark-recapture study can show a stable or growing census count while an LD-based NeN_e estimate reveals a much smaller effective size — a mismatch that is itself diagnostic, typically pointing to high variance in reproductive success (a few individuals contributing disproportionately to the next generation) or a historical bottleneck whose genetic signature outlasts the demographic recovery.

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A subscript is a label attached below a symbol. It often selects a time step, component, category, or member of a sequence.

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Sources cited in the article section

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