← All parts of this equation

Equation 22 · Part 1 · Evolutionary Biology and Ecology in Practice: An Advanced Technical Guide

Symbol hatr^2

r^2\hat{r}^2
r^2\hat{r}^2

What this part means

the mean squared correlation of allele frequencies across pairs of loci observed in the sample.

Its job in the formula

hatr2r^2 is squared: multiply the underlying quantity by itself. The square is a mathematical operation, not a second independent variable.

Where the article explains it

where r^2\hat{r}^2 is the mean squared correlation of allele frequencies across pairs of loci observed in the sample, and S is the sample size, with the 1/S term correcting for the sampling bias that a finite number of genotyped individuals introduces into r^2\hat{r}^2 even when the true population NeN_e is large.

The passage around this formula

where r^2\hat{r}^2 is the mean squared correlation of allele frequencies across pairs of loci observed in the sample, and S is the sample size, with the 1/S term correcting for the sampling bias that a finite number of genotyped individuals introduces into r^2\hat{r}^2 even when the true population NeN_e is…

Read this part in the article →

Learn the underlying idea

An exponent tells how a base is used in multiplication. In x³, x is the base and 3 is the exponent: x³ = x × x × x.

Open the illustrated exponents: repeated multiplication and powers guide →

See this notation across published equations →

Sources cited in the article section

These citations provide research context; check each source for the exact claim it supports.