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Equation 18 · Part 1 · Evolutionary Biology and Ecology in Practice: An Advanced Technical Guide

Symbol N_e

NeN_e
NeN_e

What this part means

the downward in inferred.

Its job in the formula

NeN_e is a part of this expression. Its role is fixed by the surrounding article and by the operations shown in the formula.

Where the article explains it

Physically linked loci must be excluded or flagged , because the method’s entire logic depends on treating background disequilibrium as drift-generated between unlinked loci; loci close together on the same chromosome carry disequilibrium from physical linkage that has nothing to do with NeN_e and will bias the estimate upward in apparent LD, downward in inferred NeN_e .

The passage around this formula

Effective population size, NeN_e , is not census size. It is the size of an idealized population — with random mating, no selection, non-overlapping generations, and equal sex ratio — that would show the same rate of genetic drift as the real population under study. Because NeN_e governs how fast a population loses genetic variation and how strongly drift competes with selection, it is one of the central quantities in conservation genetics: a population with a small NeN_e loses adaptive variation and accumulates inbreeding depression faster than its census size alone would suggest.

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Sources cited in the article section

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