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Equation 2 · Einstein's Random Walk and the Mathematics of Genetic Drift

What does this equation mean?

∂f(x,t)∂t=D ∂2f(x,t)∂x2\frac{\partial f(x,t)}{\partial t} = D\,\frac{\partial^{2} f(x,t)}{\partial x^{2}}

Read the formula alongside the article passage below. Each part has a deeper page with its role in the equation, the supporting passage and nearby citations.

Start withpartial^2 f(x,t)
Divide bypartial x^2
This relates tofracpartial f(x,t)partial t
How to read the two sides of this formula. Follow the article passage for the meaning of each quantity.

This equation states an equality: the expressions on both sides have the same value under the article’s assumptions. Read the equation part by part below; each part has a contextual explanation and a link to its mathematical background.

Read it piece by piece

ff

Symbol f

f occurs above the fraction bar. The numerator is divided by the entire denominator below it.

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xx

Symbol x

x is an argument of the function-like quantity on the left; its role is set by that function’s stated inputs.

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tt

Symbol t

t is an argument of the function-like quantity on the left; its role is set by that function’s stated inputs.

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DD

Symbol D

D is an input to the expression that computes the quantity on the left.

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x2x^{2}

Symbol x^2

The square of x: multiply x by itself.

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=

=

The expressions on both sides represent the same quantity under the stated assumptions.

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fraction

fraction

Divide the expression above the line by the one below it.

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derivative

derivative

This notation tracks how one quantity changes with another. The indicated variable tells which change is being measured.

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superscript

superscript

A raised number can be a power. When it is a label or bound, it selects a case or the upper limit of a sum; the formula’s structure distinguishes these uses.

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∂f(x,t)\partial f(x,t)

Numerator: partial f(x,t)

The complete quantity above the fraction bar.

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∂t\partial t

Denominator: partial t

The complete quantity below the fraction bar; it must be nonzero for this division.

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∂2f(x,t)\partial^{2} f(x,t)

Numerator: partial^2 f(x,t)

The complete quantity above the fraction bar.

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∂x2\partial x^{2}

Denominator: partial x^2

The complete quantity below the fraction bar; it must be nonzero for this division.

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How to interpret it

With a fixed numerator, increasing a nonzero denominator reduces the fraction. Read it with the definitions, units, and assumptions supplied by the article.

What the article says around this equation

Working through the statistical mechanics of a wall permeable to solvent but not to suspended particles, Einstein arrived at a differential equation for how the concentration f(x,t) of particles along one axis changes over time: ∂f(x,t)∂t=D ∂2f(x,t)∂x2\frac{\partial f(x,t)}{\partial t} = D\,\frac{\partial^{2} f(x,t)}{\partial x^{2}}. the diffusion equation, in a form still written the same way today [ 1 ] . Solving it for particles released from a point gives a spreading distribution whose variance grows linearly, not with the square root of time as a naive guess might suggest but as

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Sources cited in the surrounding passage

These citations give research context. Read each source to check which claims it supports.

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