Equation 11 · Einstein's Random Walk and the Mathematics of Genetic Drift
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Symbol D
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where R is the gas constant, T the absolute temperature, and N Avogadro’s number [ 1 ] . That last dependency was the trap door: if D could be read off a microscope by timing how far a particle wandered, and every other term in the equation was already known independently, then watching one particle jiggle became a way to count atoms.
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