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Equation 5 · Comparing the Main Approaches to Evolutionary Biology and Ecology

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ss

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The equation exposes the model’s real content: selection is strongest while the favored allele is at intermediate frequency, where pt(1−pt)p_t(1-p_t) is largest, and weakens automatically as the allele nears fixation or loss, purely from the algebra of the denominator — a pattern of decelerating change that shows up empirically in both the Grants’ beak-size tracking and the LTEE’s fitness trajectories, even though the two systems differ in essentially everything else about how the selection coefficient s itself is generated.

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