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Equation 1 · Part 2 · Why We Punish Strangers: The Evolution of Human Cooperation

Symbol b

rb>crb > c
bb

What this part means

the fitness benefit the recipient gains.

Its job in the formula

b is a part of this expression. Its role is fixed by the surrounding article and by the operations shown in the formula.

Where the article explains it

Here r is the coefficient of relatedness between the actor and the recipient, b is the fitness benefit the recipient gains, and c is the fitness cost the actor pays.

The passage around this formula

…of social behaviour gave altruism toward kin a precise condition under which natural selection favors it [ 1 ] : rb>crb > c. Here r is the coefficient of relatedness between the actor and the recipient, b is the fitness benefit the recipient gains, and c is the fitness cost the actor pays. A gene that disposes its carrier to help a relative will spread whenever the benefit to the recipient, discounted by how likely the recipient is to carry a copy of the same gene, exceeds the cost to the helper.…

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A variable is a named place for a value. Its letter is a local label: x can mean position in one formula and a data point in another.

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Sources cited in the surrounding passage

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