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Equation 9 · Part 10 · The Peacock Problem: Darwin's Second Theory and Its Hard Tests

addition

Δtˉ=ht2 βt+B βpΔpˉ=B βt+hp2 βp\begin{aligned} \Delta \bar{t} &= h_t^{2}\,\beta_t + B\,\beta_p \\ \Delta \bar{p} &= B\,\beta_t + h_p^{2}\,\beta_p \end{aligned}
addition

What this part means

Add the term after the plus sign to the term or group before it.

Its job in the formula

Add the term after the plus sign to the term or group before it.

The passage around this formula

Russell Lande’s 1981 paper gave this verbal logic its standard quantitative-genetic form, modelling the joint evolution of a female preference trait and a male display trait under stabilising natural selection on the display and showed that, “despite stabilizing natural selection on males, various types of mating preferences may create a runaway process in which the outcome of phenotypic evolution depends critically on the genetic variation parameters and initial conditions of a population” [ 4 ] . One compact way to see why sits in the coupling itself. Let tˉ\bar t be the population mean of a male display trait and pˉ\bar p the population mean of the corresponding female preference; write…

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Learn the underlying idea

Addition combines quantities; subtraction measures the signed difference between them. Parentheses show what is combined before the rest of the expression is evaluated.

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Sources cited in the surrounding passage

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