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Equation 7 · Part 1 · The Peacock Problem: Darwin's Second Theory and Its Hard Tests

Symbol h_p^2

hp2h_p^2
hp2h_p^2

What this part means

hp2h_p^2 is squared: multiply the underlying quantity by itself. The square is a mathematical operation, not a second independent variable.

Its job in the formula

hp2h_p^2 is squared: multiply the underlying quantity by itself. The square is a mathematical operation, not a second independent variable.

The passage around this formula

…population mean of the corresponding female preference; write βt\beta_t and βp\beta_p for the direct selection gradients acting on each (natural selection pulling βt\beta_t downward, since the ornament is costly), ht2h_t^2 and hp2h_p^2 for their heritabilities, and B for the genetic covariance that non-random mating builds between the two traits. The standard Lande-Fisher formalisation of one generation’s change is then a coupled pair:

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A subscript is a label attached below a symbol. It often selects a time step, component, category, or member of a sequence.

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Sources cited in the surrounding passage

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