Symbol beta_t

Its job in the formula

bet$a_t$ is a part of this expression. Its role is fixed by the surrounding article and by the operations shown in the formula.

Where the article explains it

Let $\bar t$ be the population mean of a male display trait and $\bar p$ the population mean of the corresponding female preference; write $\beta_t$ and $\beta_p$ for the direct selection gradients acting on each (natural selection pulling $\beta_t$ downward, since the ornament is costly), $h_t^2$ and $h_p^2$ for their heritabilities, and B for the genetic covariance that non-random mating builds between the two traits.

Learn the underlying idea

A subscript is a label attached below a symbol. It often selects a time step, component, category, or member of a sequence.

Open the illustrated subscripts: which member of a family? guide →