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Equation 1 · From Origins to Frontier: A History of Evolutionary Biology and Ecology

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NeN_e

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NeN_e

Symbol N_e

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subscript

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Sewall Wright’s 1931 paper, “Evolution in Mendelian Populations,” added the piece Fisher’s and Haldane’s more deterministic treatments lacked: a formal account of genetic drift, the random change in allele frequency from generation to generation caused by finite population size [ 3 ] . Wright showed that in a randomly mating population of effective size NeN_e , the variance in allele frequency change per generation from sampling alone is

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