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Equation 2 · Part 1 · Darwin's Finches: The Beak the Grants Measured for Forty Years

Symbol S

SS
SS

What this part means

the selection differential actually measured on the population in one generation, h2h^2 is the trait’s heritability.

Its job in the formula

S is a part of this expression. Its role is fixed by the surrounding article and by the operations shown in the formula.

Where the article explains it

where S is the selection differential actually measured on the population in one generation, h2h^2 is the trait’s heritability, and R is the response: the amount the population’s mean should shift in the next generation, if the difference between survivors and non-survivors was genetic rather than purely environmental.

The passage around this formula

where S is the selection differential actually measured on the population in one generation, h2h^2 is the trait’s heritability, and R is the response: the amount the population’s mean should shift in the next generation, if the difference between survivors and non-survivors was genetic rather than purely…

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Sources cited in the surrounding passage

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