Equation 1 · Part 4 · Darwin's Finches: The Beak the Grants Measured for Forty Years
=
=
What this part means
The expressions on both sides represent the same quantity under the stated assumptions.
Its job in the formula
The equals sign connects the complete expression on the left with the complete expression on the right. Both sides must have compatible units.
Full expression→=→Article meaning
The passage around this formula
Heritability is what turns a measured selection differential into a testable prediction, through the simplest quantitative-genetics identity in the field, the breeder’s equation: . where S is the selection differential actually measured on the population in one generation, is the trait’s heritability, and R is the response: the amount the population’s mean should shift in the next generation, if the difference between survivors and non-survivors was genetic rather than purely environmental. Grant and Grant ran exactly this check on their own data, comparing the mean of a trait before a selection episode with the mean of the same trait in the following generation. The…
Learn the underlying idea
An equals sign says that the expression on its left and the expression on its right have the same value under the stated definitions and assumptions.
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Sources cited in the surrounding passage
These citations provide research context; check each source for the exact claim it supports.